(Shk4_Ddis:0.3241886452,Shk5_Ddis:0.5473223828,(Shk2_Ddis:1.1330540142,(Shk1_Ddis:0.3450754868,Shk3_Ddis:0.4528002100)80:0.0494350228)96:0.2278505623);