(CR4L_e-1_S:0.0000000001,CR4L_e-2_S:0.0222616933,(CR4L_d_Smo:1.3070808620,((CR4L_a-1_S:0.0024021510,CR4L_a-2_S:0.0000016203)100:0.5789635364,(CR4L_b_Smo:0.2036162394,CR4L_c_Smo:0.1164775098)100:1.0734389877)100:0.5418771275)100:1.0665267887);